Research Article

Abstract
References
1

Aakash A, Thakur NS, Singh MK, Bhayal L, Meena K, Choudhary SK, Kumawat N, Singh RK, Singh UP, et al. (2022) Sustainability in rainfed maize (Zea mays L.) production using choice of corn variety and nitrogen scheduling. Sustainability 14:3116. https://doi.org/10.3390/su14053116

10.3390/su14053116
2

Abbott JA (1999) Quality measurement of fruits and vegetables. Postharvest Biol Technol 15:207-225. https://doi.org/10.1016/S0925-5214(98)00086-6

10.1016/S0925-5214(98)00086-6
3

Ahn JJ, Kim EA, Shin EJ, Sunwoo Y, Lee JH, Nam SY (2024) Shading treatments affect the growth characteristics, ornamental value, and photosynthetic activities of various Peperomia species and cultivars. Glasilo Future 7:1-19. https://doi.org/10.32779/gf.7.2-3.1

10.32779/gf.7.2-3.1
4

Alcántara-Ayala O, Oyama K, Ríos-Muñoz CA, Rivas G, Ramirez-Barahona S, Luna-Vega I (2020) Morphological variation of leaf traits in the Ternstroemia lineata species complex (Ericales: Penthaphylacaceae) in response to geographic and climatic variation. PeerJ 8:e8307. https://doi.org/10.7717/peerj.8307

10.7717/peerj.830731942256PMC6956789
5

Alonso RS, Sodré GA, Silva DC (2024) Dickson quality index of cocoa genotypes under water deficit. Forests 15:2054. https://doi.org/10.3390/f15122054

10.3390/f15122054
6

Altesor A, Ezcurra E (2003) Functional morphology and evolution of stem succulence in cacti. J Arid Environ 53:557-567. https://doi.org/10.1006/jare.2002.1059

10.1006/jare.2002.1059
7

Amzallag GN (2001) Data analysis in plant physiology: are we missing the reality? Plant Cell Environ 24:881-890. https://doi.org/10.1046/j.1365-3040.2001.00742.x

10.1046/j.1365-3040.2001.00742.x
8

Bae YH, Yang HC, Kim YH, Hyeon SJ, Choi MS, Vu NT, Jang DC (2025) Seedling quality and early growth after the transplanting of tomato seedlings grown using different production methods. Hortic Sci Technol 43:275-285. https://doi.org/10.7235/HORT.20250025

10.7235/HORT.20250025
9

Baginsky S, Hennig L, Zimmermann P, Gruissem W (2010) Gene expression analysis, proteomics, and network discovery. Plant Physiol 152:402-410. https://doi.org/10.1104/pp.109.150433

10.1104/pp.109.15043320018595PMC2815903
10

Bai Y, Shi W, Xing X, Wang Y, Jin Y, Zhang L, Song YF, Dong LH, Liu HB (2014) Study on tobacco vigorous seedling indexes model. Sci Agric Sin 6:1086-1098. https://doi.org/10.3864/j.issn.0578-1752.2014.06.005

10.3864/j.issn.0578-1752.2014.06.005
11

Bannari A, Morin D, Bonn F, Huete A (1995) A review of vegetation indices. Remote Sen Rev 13:95-120. https://doi.org/10.1080/02757259509532298

10.1080/02757259509532298
12

Bayala J, Dianda M, Wilson J, Ouedraogo SJ, Sanon K (2009) Predicting field performance of five irrigated tree species using seedling quality assessment in Burkina Faso, West Africa. New For 38:309-322. https://doi.org/10.1007/s11056-009-9149-4

10.1007/s11056-009-9149-4
13

Bhagsari AS, Brown RH (1986) Leaf photosynthesis and its correlation with leaf area. Crop Sci 26:127-132. https://doi.org/10.2135/cropsci1986.0011183X002600010030x

10.2135/cropsci1986.0011183X002600010030x
14

Binotto AF, Lúcio ADC, Lopes SJ (2010) Correlations between growth variables and the Dickson quality index in forest seedlings. Cerne 16:457-464. https://doi.org/10.1590/S0104-77602010000400005

10.1590/S0104-77602010000400005
15

Bláha L (2021) Importance of root-shoot ratio for crops production: a review. Curr Top Agric Sci 1:37-49. https://doi.org/10.9734/bpi/ctas/v1/12112D

10.9734/bpi/ctas/v1/12112D
16

Boke NH (1980) Developmental morphology and anatomy in Cactaceae. BioScience 30:605-610. https://doi.org/10.2307/1308111

10.2307/1308111
17

Brisco B, Brown RJ, Hirose T, McNairn H, Staenz K (1998) Precision agriculture and the role of remote sensing: a review. Can J Remote Sens 24:315-327. https://doi.org/10.1080/07038992.1998.10855254

10.1080/07038992.1998.10855254
18

Cabahug RAM, Nam SY, Lim KB, Jeon JK, Hwang YJ (2018) Propagation techniques for ornamental succulents. Flower Res J 26:90-101. https://doi.org/10.11623/frj.2018.26.3.02

10.11623/frj.2018.26.3.02
19

Choi EJ, Kim JY, Jang GH, Kim SH, Oh SI, Koh HM, Hong S, Yoon M, Ko CH (2024a) Seedling growth of Lysimachia mauritiana Lam. by cell size and nutrient solution concentration. Flower Res J 32:103-109. https://doi.org/10.11623/frj.2024.32.2.05

10.11623/frj.2024.32.2.05
20

Choi HE, Hwang SY, Yun JH, Yu J, Hwang JH, Park EW, Koo JK, Hwang SJ (2024b) Growth of grafted cucumber seedlings as affected by different EC levels and cultivars. J Bio Environ Control 33:387-396. https://doi.org/10.12791/KSBEC.2024.33.4.387

10.12791/KSBEC.2024.33.4.387
21

Cochavi A, Cohen IH, Rachmilevitch S (2020) The role of different root orders in nutrient uptake. Environ Exp Bot 179:104212. https://doi.org/10.1016/j.envexpbot.2020.104212

10.1016/j.envexpbot.2020.104212
22

Cohen MN (2009) Introduction: rethinking the origins of agriculture. Curr Anthropol 50:591-595. https://doi.org/10.1086/603548

10.1086/603548
23

Currey CJ, Torres AP, Lopez RG, Jacobs DF (2011) The quality index-a new tool for integrating quantitative measurements to assess quality of young floriculture plants. Acta Hortic 1000:385-391. https://doi.org/10.17660/ActaHortic.2013.1000.53

10.17660/ActaHortic.2013.1000.53
24

De Vylder J, Vandenbussche F, Hu Y, Philips W, Van Der Straeten D (2012) Rosette tracker: an open source image analysis tool for automatic quantification of genotype effects. Plant Physiol 160:1149-1159. https://doi.org/10.1104/pp.112.202762

10.1104/pp.112.20276222942389PMC3490612
25

Díaz S, Cabido M (2001) Vive la différence: plant functional diversity matters to ecosystem processes. Trends Ecol Evol 16:646-655. https://doi.org/10.1016/S0169-5347(01)02283-2

10.1016/S0169-5347(01)02283-2
26

Dickson A, Leaf AL, Hosner JF (1960) Quality appraisal of white spruce and white pine seedling stock in nurseries. For Chron 36:10-13. https://doi.org/10.5558/tfc36010-1

10.5558/tfc36010-1
27

Dorsey BL, Haevermans T, Aubriot X, Morawetz JJ, Riina R, Steinmann VW, Berry PE (2013) Phylogenetics, morphological evolution, and classification of Euphorbia subgenus Euphorbia. Taxon 62:291-315. https://doi.org/10.12705/622.1

10.12705/622.1
28

Dos Santos P, Brilhante MÂ, Messerschmid TF, Serrano HC, Kadereit G, Branquinho C, De Vos JM (2022) Plant growth forms dictate adaptations to the local climate. Front Plant Sci 13:1023595. https://doi.org/10.3389/fpls.2022.1023595

10.3389/fpls.2022.102359536479511PMC9720395
29

Duncan WG, Hesketh JD (1968) Net photosynthetic rates, relative leaf growth rates, and leaf numbers of 22 races of maize grown at eight temperatures. Crop Sci 8:670-674. https://doi.org/10.2135/cropsci1968.0011183X000800060009x

10.2135/cropsci1968.0011183X000800060009x
30

Egerton FN (2008) A history of the ecological sciences, part 28: plant growth studies during the 1700s. Bull Ecol Soc Am 89:159-175.

10.1890/0012-9623(2008)89[159:AHOTES]2.0.CO;2
31

Elgaml NM, Salama AB, Shehata HS, Abdelhamid MT (2022) Effective microorganisms improve growth, nutrients uptake, normalized difference vegetation index, photosystem II, and essential oil while reducing canopy temperature in water-stressed Salvia sclarea plants. Int J Agron 2022:1767347. https://doi.org/10.1155/2022/1767347

10.1155/2022/1767347
32

El-Sharkawy MA (2011) Overview: early history of crop growth and photosynthesis modeling. BioSystems 103:205-211. https://doi.org/10.1016/j.biosystems.2010.08.004

10.1016/j.biosystems.2010.08.004
33

Fleisher DH, Timlin DJ, Reddy VR (2006) Temperature influence on potato leaf and branch distribution and on canopy photosynthetic rate. Agron J 98:1442-1452. https://doi.org/10.2134/agronj2005.0322

10.2134/agronj2005.0322
34

Freschet GT, Swart EM, Cornelissen JH (2015) Integrated plant phenotypic responses to contrasting above- and below-ground resources: key roles of specific leaf area and root mass fraction. New Phytol 206:1247-1260. https://doi.org/10.1111/nph.13352

10.1111/nph.13352
35

Gallegos-Cedillo VM, Diánez F, Nájera C, Santos M (2021) Plant agronomic features can predict quality and field performance: a bibliometric analysis. Agronomy 11:2305. https://doi.org/10.3390/agronomy11112305

10.3390/agronomy11112305
36

Garty J, Weissman L, Tamir O, Beer S, Cohen Y, Karnieli A, Orlovsky L (2000) Comparison of five physiological parameters to assess the vitality of the lichen Ramalina lacera exposed to air pollution. Physiol Plant 109:410-418. https://doi.org/10.1034/j.1399-3054.2000.100407.x

10.1034/j.1399-3054.2000.100407.x
37

Go Y, Ju SH, Kim EJ, Kim YJ, Lee BS, Na H (2025) Growth and quality of grafted watermelon seedlings under controlled light intensity in a plant factory nursery system. J Bio Environ Control 34:154-161. https://doi.org/10.12791/KSBEC.2025.34.2.154

10.12791/KSBEC.2025.34.2.154
38

Govaerts B, Verhulst N (2010) The normalized difference vegetation index (NDVI) greenseeker (TM) handheld sensor: toward the integrated evaluation of crop management. Part A: concepts and case studies. International Maize and Wheat Improvement Center (CIMMYT), Texcoco, Mexico

39

Griffiths H, Males J (2017) Succulent plants. Curr Biol 27:R890-R896. https://doi.org/10.1016/j.cub.2017.03.021

10.1016/j.cub.2017.03.021
40

Grossnickle SC (2012) Why seedlings survive: influence of plant attributes. New For 43:711-738. https://doi.org/10.1007/s11056-012-9336-6

10.1007/s11056-012-9336-6
41

Güney D, Bayraktar A, Atar F, Turna İ (2020) Effects of root undercutting, fertilization and thinning on seedling growth and quality of oriental beech (Fagus orientalis Lipsky) seedlings. Artvin Çoruh Univ Orman Fak Derg 21:214-222. https://doi.org/10.17474/artvinofd.696079

10.17474/artvinofd.696079
42

Guo Y, Guo H, Liu Y, Tao Y, Liang Y, Gao Z, Tang K, Feng F (2024) Combination of silicon and plant growth promoting rhizobacteria consortia promoted the growth of melon seedlings under salt stress. Hortic Environ Biotechnol 65:1069-1078. https://doi.org/10.1007/s13580-024-00633-3

10.1007/s13580-024-00633-3
43

Ha STT, Kim YT, In BC (2025) Pre and postharvest application of nano silver improves gray mold disease resistance and longevity of cut roses. Hortic Environ Biotechnol 66:173-187. https://doi.org/10.1007/s13580-024-00647-x

10.1007/s13580-024-00647-x
44

Haworth M, Marino G, Atzori G, Fabbri A, Daccache A, Killi D, Carli A, Montesano V, Conte A, et al. (2023) Plant physiological analysis to overcome limitations to plant phenotyping. Plants 12:4015. https://doi.org/10.3390/plants12234015

10.3390/plants1223401538068650PMC10708318
45

Hilty J, Muller B, Pantin F, Leuzinger S (2021) Plant growth: the what, the how, and the why. New Phytol 232:25-41. https://doi.org/10.1111/nph.17610

10.1111/nph.17610
46

Hoang NH, Kane ME, Radcliffe EN, Zettler LW, Richardson LW (2017) Comparative seed germination and seedling development of the ghost orchid, Dendrophylax lindenii (Orchidaceae), and molecular identification of its mycorrhizal fungus from South Florida. Ann Bot 119:379-393. https://doi.org/10.1093/aob/mcw220

10.1093/aob/mcw22028025292PMC5314644
47

Holmes GD (1975) History of forestry and forest management. Philos Trans R Soc Lond B Biol Sci 271:69-80. https://doi.org/10.1098/rstb.1975.0035

10.1098/rstb.1975.0035
48

Hong J, Lee JH, Kim EA, Shin EJ, Lee S, Joo S, Cha EB, Han H, Lee EC, et al. (2025) Dry-cold stratification combined with pericarp removal: germination, seedling development, and physiological changes of lilyturf (Liriope muscari (Decne.) L.H. Bailey). J People Plants Environ 28:777-798. https://doi.org/10.11628/ksppe.2025.28.6.777

10.11628/ksppe.2025.28.6.777
49

Huang S, Tang L, Hupy JP, Wang Y, Shao G (2021) A commentary review on the use of normalized difference vegetation index (NDVI) in the era of popular remote sensing. J For Res 32:1-6. https://doi.org/10.1007/s11676-020-01155-1

10.1007/s11676-020-01155-1
50

Hunt R, Causton DR, Shipley B, Askew AP (2002) A modern tool for classical plant growth analysis. Ann Bot 90:485-488. https://doi.org/10.1093/aob/mcf214

10.1093/aob/mcf21412324272PMC4240380
51

Hwang JH, Park EW, Koo JK, Hwang HS, Yu J, Yun JH, Hwang SY, Choi HE, Yoon YH, et al. (2025) Controlling plant height and internode length in balloon flower and lance asiabell seedling using different light qualities. J Bio Environ Control 34:207-219. https://doi.org/10.12791/KSBEC.2025.34.2.207

10.12791/KSBEC.2025.34.2.207
52

Iacobucci D, Román S, Moon S, Rouziès D (2025) A tutorial on what to do with skewness, kurtosis, and outliers: new insights to help scholars conduct and defend their research. Psychol Mark 42:1398-1414. https://doi.org/10.1002/mar.22187

10.1002/mar.22187
53

Ibaraki Y, Murakami J (2006) Distribution of chlorophyll fluorescence parameter Fv/Fm within individual plants under various stress conditions. Acta Hortic 761:255-260. https://doi.org/10.17660/ActaHortic.2007.761.33

10.17660/ActaHortic.2007.761.33
54

Ingestad T, Agren GI (1991) The influence of plant nutrition on biomass allocation. Ecol Appl 1:168-174. https://doi.org/10.2307/1941809

10.2307/1941809
55

Inthima P, Supaibulwatana K (2025) Green LEDs lighting enhances vigorous growth and boosts bacoside production in hydroponically cultivated Bacopa monnieri (L.) Wettst. Hortic Environ Biotechnol 66:491-507. https://doi.org/10.1007/s13580-024-00666-8

10.1007/s13580-024-00666-8
56

Jang HS, Kang YI, Suh JN, Ahn MS, Jung HH, Oh SI, Seo KH, Yi PH (2025) Assessment of drought tolerance for herbaceous plants for low-maintenance roadside flowerscape. Flower Res J 33:83-92. https://doi.org/10.11623/frj.2025.33.2.05

10.11623/frj.2025.33.2.05
57

Jang IT, Lee JH, Shin EJ, Nam SY (2023) Evaluation of growth, flowering, and chlorophyll fluorescence responses of Viola cornuta cv. Penny Red Wing according to spectral power distributions. J People Plants Environ 26:335-349. https://doi.org/10.11628/ksppe.2023.26.4.335

10.11628/ksppe.2023.26.4.335
58

Jaskowiak PA, Campello RJ, Costa IG (2014) On the selection of appropriate distances for gene expression data clustering. BMC Bioinformatics 15:S2. https://doi.org/10.1186/1471-2105-15-S2-S2

10.1186/1471-2105-15-S2-S224564555PMC4072854
59

Jeong HW, Lee HR, Kim HM, Kim HM, Hwang HS, Hwang SJ (2020) Using light quality for growth control of cucumber seedlings in closed-type plant production system. Plants 9:639. https://doi.org/10.3390/plants9050639

10.3390/plants905063932429540PMC7284851
60

Jo S, Kim DH, Yoon JW, Cheong EJ (2025) Shading treatment-induced changes with growth and physiological characteristics in Delphinium maackianum Regel. Korean J Plant Resour 38:300-312. https://doi.org/10.7732/kjpr.2025.38.3.300

10.7732/kjpr.2025.38.3.300
61

Jolliffe PA, Eaton GW, Doust JL (1982) Sequential analysis of plant growth. New Phytol 92:287-296. https://doi.org/10.1111/j.1469-8137.1982.tb03387.x

10.1111/j.1469-8137.1982.tb03387.x
62

Jones G, Kluyver T, Preece C, Swarbrick J, Forster E, Wallace M, Charles M, Rees M, Osborne CP (2021) The origins of agriculture: intentions and consequences. J Archaeol Sci 125:105290. https://doi.org/10.1016/j.jas.2020.105290

10.1016/j.jas.2020.105290
63

Khan S, Purohit A, Vadsaria N (2020) Hydroponics: current and future state of the art in farming. J Plant Nutr 44:1515-1538. https://doi.org/10.1080/01904167.2020.1860217

10.1080/01904167.2020.1860217
64

Khanh HTTM, Cabahug-Braza RAM, Lim KB, Hwang YJ (2023) A review: leaf anatomy features of Echeveria. Flower Res J 31:1-9. https://doi.org/10.11623/frj.2023.31.1.01

10.11623/frj.2023.31.1.01
65

Khanh HTTM, Cabahug-Braza RAM, Lim KB, Hwang YJ (2024) Leaf morpho-anatomical evaluation of Echeveria cultivars using different fixation methods. Hortic Sci Technol 42:127-142. https://doi.org/10.7235/HORT.20240011

10.7235/HORT.20240011
66

Kim D, Goo H, Yoon S, Yoon J, Kim H, Yoo Y, Lee M, Park KS (2025a) Effects of supplemental LED lighting based on solar irradiation and carbon dioxide enrichment on photosynthesis, growth, and yield of greenhouse-grown tomato plants. J People Plants Environ 28:1-12. https://doi.org/10.11628/ksppe.2025.28.1.1

10.11628/ksppe.2025.28.1.1
67

Kim EA, Lee JH, Nam SY (2024a) Differences in growth characteristics and photochemical responses of Salvia miltiorrhiza Bunge under different LED light qualities in a closed plant factory system. Korean J Med Crop Sci 32:189-201. https://doi.org/10.7783/KJMCS.2024.32.4.189

10.7783/KJMCS.2024.32.4.189
68

Kim EA, Lee JH, Nam SY (2024b) Experimental control of day and night temperatures to improve the growth and external quality of Salvia miltiorrhiza Bunge. Hortic Sci Technol 42:94-103. https://doi.org/10.7235/HORT.20240008

10.7235/HORT.20240008
69

Kim EA, Lee JH, Shin EJ, Ahn JJ, Nam SY (2024c) Investigation of germination success, post-germination growth, and physiological characteristics of Betula platyphylla var. japonica seeds subjected to various short-term dry-cold treatment periods. J Agric Life Environ Sci 36:217-231. https://doi.org/10.22698/jales.20240018

10.22698/jales.20240018
70

Kim KH, Shawon MRA, An JH, Lee HJ, Kwon DJ, Hwang IC, Bae JH, Choi KY (2022) Effect of shade screen on sap flow, chlorophyll fluorescence, NDVI, plant growth and fruit characteristics of cultivated paprika in greenhouse. Agriculture 12:1405. https://doi.org/10.3390/agriculture12091405

10.3390/agriculture12091405
71

Kim Y, Yang H, Bae Y, Hyeon S, Choi M, Vu NT, Jang D (2025b) Effect of high concentration of KCl treatment on tomato seedling for overgrowth suppression and drought stress tolerance enhancement in plant factory with artificial lighting. Hortic Environ Biotechnol 66:1005-1014. https://doi.org/10.1007/s13580-025-00713-y

10.1007/s13580-025-00713-y
72

Kim YJ, Ju SH, Kim EJ, Hwang H, Na H (2025c) Optimal air temperature for producing high-quality grafted watermelon seedlings in a plant factory system. J Bio Environ Control 34:393-400. https://doi.org/10.12791/KSBEC.2025.34.4.393

10.12791/KSBEC.2025.34.4.393
73

Kim YJ, Yeom MS, Lee JY, Park SY, Oh MM (2024d) Biomass and ginsenosides content of ginseng sprouts upon exposure to several white LED lights with different correlated color temperatures and color rendering indexes. Hortic Sci Technol 42:167-179. https://doi.org/10.7235/HORT.20240014

10.7235/HORT.20240014
74

Ko BU, Bae JH, Hwang SJ, Kim HC (2017) Seedling qualities of watermelon as affected by different raising seedling period and growth characteristics after planting. J Bio Environ Control 26:56-63. https://doi.org/10.12791/KSBEC.2017.26.2.56

10.12791/KSBEC.2017.26.2.56
75

Korn RW (2011) Window patterns in Lithops. Int J Plant Sci 172:1101-1109. https://doi.org/10.1086/662031

10.1086/662031
76

Lee JH (2023) Effects of light quality, temperature, photoperiod, and GA3 concentration on growth of six Phedimus species. Ph.D. dissertation, Sahmyook University, Seoul, Korea.

77

Lee JH, Cabahug RAM, You NH, Nam SY (2021) Chlorophyll fluorescence and growth evaluation of ornamental foliage plants in response to light intensity levels under continuous lighting conditions. Flower Res J 29:153-164. https://doi.org/10.11623/frj.2021.29.3.05

10.11623/frj.2021.29.3.05
78

Lee JH, Jang IT, Kim EA, Shin EJ, Lee S, Lee M, Nam SY (2024a) Evaluating the influence of various light spectra on the growth and morphological responses of air plant (Tillandsia ionantha Planch.) grown under non-substrate and restricted irrigation conditions in a controlled environment facility. J Agric Life Environ Sci 36:546-561. https://doi.org/10.22698/jales.20240042

10.22698/jales.20240042
79

Lee JH, Kim EA, Sunwoo Y, Cha EB, Joo S, Lee S, Lee JG, Nam SY (2025a) Light spectrum strategies to optimize yield and physiological performance of Korean thistle (Cirsium setidens (Dunn) Nakai) in a closed-type plant production system. J People Plants Environ 28:799-823. https://doi.org/10.11628/ksppe.2025.28.6.799

10.11628/ksppe.2025.28.6.799
80

Lee JH, Lee S, Lee JG, Hwang SY, Nam SY (2025b) Photoperiod optimization in controlled environments: interspecific differences in vegetative growth and photochemical efficiency among three Phedimus species native to Korea. Korean J Plant Resour 38:826-841. https://doi.org/10.7732/kjpr.2025.38.6.826

10.7732/kjpr.2025.38.6.826
81

Lee JH, Lee S, Nam SY (2025c) Optimized concentrations of auxinic rooting promoters improve stem cutting propagation efficiency and morphophysiological characteristics in Hedera algeriensis cv. Gloire de Marengo. Hortic Sci Technol 43:357-372. https://doi.org/10.7235/HORT.20250032

10.7235/HORT.20250032
82

Lee JH, Nam SY (2023a) Comparison of growth and leaf color quality of Mesembryanthemum cordifolium f. variegata as affected by shading levels. J People Plants Environ 26:207-217. https://doi.org/10.11628/ksppe.2023.26.3.207

10.11628/ksppe.2023.26.3.207
83

Lee JH, Nam SY (2023b) Vegetative propagation of six Pachyphytum species as influenced by different LED light qualities. Hortic Sci Technol 41:237-249. https://doi.org/10.7235/HORT.20230022

10.7235/HORT.20230022
84

Lee JH, Nam SY (2024) Growth and photosynthetic responses of Taraxacum coreanum Nakai seedlings according to the shading levels. Hortic Sci Technol 42:214-224. https://doi.org/10.7235/HORT.20240018

10.7235/HORT.20240018
85

Lee JH, Shin EJ, Kim EA, Jang IT, Lee S, Nam SY (2024b) Effects of different concentrations of exogenous auxins (IAA, IBA, and NAA) on growth and rooting ability of Philodendron hederaceum var. oxycardium (Schott) Croat stem cuttings. J People Plants Environ 27:279-289. https://doi.org/10.11628/ksppe.2024.27.4.279

10.11628/ksppe.2024.27.4.279
86

Lee JH, Shin EJ, Nam SY (2023) Enhancing propagation of Orostachys spp. offsets through exogenous GA3 application. Horticulturae 9:1280. https://doi.org/10.3390/horticulturae9121280

10.3390/horticulturae9121280
87

Lim YJ, Kwak MJ, Lee J, Kang D, Je SM, Woo SY (2024) Korean flowering cherry (Prunus × yedoensis Matsum.) response to elevated ozone: physiological traits and biogenic volatile organic compounds emission. Hortic Environ Biotechnol 65:1025-1042. https://doi.org/10.1007/s13580-024-00628-0

10.1007/s13580-024-00628-0
88

Lin KH, Wu CW, Chang YS (2019) Applying Dickson quality index, chlorophyll fluorescence, and leaf area index for assessing plant quality of Pentas lanceolata. Not Bot Horti Agrobot Cluj Napoca 47:169-176. https://doi.org/10.15835/nbha47111312

10.15835/nbha47111312
89

Liu X, Shi R, Gao M, He R, Li Y, Liu H (2023) Growth of tomato and cucumber seedlings under different light environments and their development after transplanting. Front Plant Sci 14:1164768. https://doi.org/10.3389/fpls.2023.1164768

10.3389/fpls.2023.116476837546262PMC10400448
90

Lozano-Castellanos LF, Navas-Gracia LM, Lozano-Castellanos IC, Correa-Guimaraes A (2025) Technologies applied to artificial lighting in indoor agriculture: a review. Sustainability 17:3196. https://doi.org/10.3390/su17073196

10.3390/su17073196
91

Mohi-Ud-Din M, Hossain MA, Rohman MM, Uddin MN, Haque MS, Ahmed JU, Hossain A, Hassan MM, Mostofa MG (2021) Multivariate analysis of morpho-physiological traits reveals differential drought tolerance potential of bread wheat genotypes at the seedling stage. Plants 10:879. https://doi.org/10.3390/plants10050879

10.3390/plants1005087933925375PMC8145240
92

Morales-Gallegos LM, Martínez-Trinidad T, Hernández-de la Rosa P, Gómez-Guerrero A, Alvarado-Rosales D, Saavedra-Romero LDL (2023) Tree health condition in urban green areas assessed through crown indicators and vegetation indices. Forests 14:1673. https://doi.org/10.3390/f14081673

10.3390/f14081673
93

Mota CS, Araújo ELS, Silva FG, Dornelles P, Freiberger MB, Mendes GC (2018) Physiology and quality of Eugenia dysenterica DC seedlings grown in vermiculite and rice husk-based substrates. Rev Bras Frutic 40:e-049. https://doi.org/10.1590/0100-29452018049

10.1590/0100-29452018049
94

Muniz CR, Freire FCO, Viana FMP, Cardoso JE, Sousa CAF, Guedes MIF, van der Schoor R, Jalink H (2014) Monitoring cashew seedlings during interactions with the fungus Lasiodiplodia theobromae using chlorophyll fluorescence imaging. Photosynthetica 52:529-537. https://doi.org/10.1007/s11099-014-0061-6

10.1007/s11099-014-0061-6
95

Musa FI, Sahoo UK (2025) Impact of pre-treatments on Albizia procera and Albizia chinensis seed germination and early growth performance in nursery, Mizoram, India. For Sci Technol 21:138-148. https://doi.org/10.1080/21580103.2025.2465604

10.1080/21580103.2025.2465604
96

Nam HH, Kim KS, Kim BS (2022) Seed harvesting and seedling raising methods for seedling production in Angelica dahurica. Korean J Med Crop Sci 30:180-186. https://doi.org/10.7783/KJMCS.2022.30.3.180

10.7783/KJMCS.2022.30.3.180
97

Nam SY (2024) Succulent plant. RGB press, Seoul, Korea

98

Niinemets Ü (2007) Photosynthesis and resource distribution through plant canopies. Plant Cell Environ 30:1052-1071. https://doi.org/10.1111/j.1365-3040.2007.01683.x

10.1111/j.1365-3040.2007.01683.x
99

Niinemets Ü, Tobias M, Cescatti A, Sparrow A (2006) Size-dependent variation in shoot light-harvesting efficiency in shade-intolerant conifers. Int J Plant Sci 167:19-32. https://doi.org/10.1086/497845

10.1086/497845
100

Niklas KJ (1994) Plant allometry: the scaling of form and process. University of Chicago Press, Chicago, USA.

101

Nyoka BI, Kamanga R, Njoloma J, Jamnadass R, Mng’omba S, Muwanje S (2018) Quality of tree seedlings produced in nurseries in Malawi: an assessment of morphological attributes. For Trees Livelihoods 27:103-117. https://doi.org/10.1080/14728028.2018.1443027

10.1080/14728028.2018.1443027
102

Opel MR (2005) A morphological phylogeny of the genus Conophytum NE Br. (Aizoaceae). Haseltonia 2005:53-77. https://doi.org/10.2985/1070-0048(2005)11[53:AMPOTG]2.0.CO;2

10.2985/1070-0048(2005)11[53:AMPOTG]2.0.CO;2
103

Park BG, Lee JH, Shin EJ, Kim EA, Nam SY (2024) Light quality influence on growth performance and physiological activity of Coleus cultivars. Int J Plant Biol 15:807-826. https://doi.org/10.3390/ijpb15030058

10.3390/ijpb15030058
104

Park EW, Hwang JH, Hwang HS, Jeong HW, Hwang SY, Yu J, Hwang SJ (2023a) Appropriate cold treatment periods and shading levels on Codonopsis lanceolata for plug seedling production in summer season. J Bio Environ Control 32:157-164. https://doi.org/10.12791/KSBEC.2023.32.2.157

10.12791/KSBEC.2023.32.2.157
105

Park SH, Lee JH, Nam SY (2023b) An analysis of the growth and photosynthetic responses of potted Veronica pusanensis Y.N.Lee according to the shading levels. J People Plants Environ 26:219-231. https://doi.org/10.11628/ksppe.2023.26.3.219

10.11628/ksppe.2023.26.3.219
106

Park SH, Park HG, Kwon HH, Lee KM (2023c) Assessment of adequate volumetric water content for managing Farfugium japonicum (L.) Kitam. in the indoor gardens. J People Plants Environ 26:749-759. https://doi.org/10.11628/ksppe.2023.26.6.749

10.11628/ksppe.2023.26.6.749
107

Pettorelli N (2013) The normalized difference vegetation index. Oxford University Press, New York, USA.

10.1093/acprof:osobl/9780199693160.001.0001
108

Pettorelli N, Ryan S, Mueller T, Bunnefeld N, Jędrzejewska B, Lima M, Kausrud K (2011) The normalized difference vegetation index (NDVI): unforeseen successes in animal ecology. Clim Res 46:15-27. https://doi.org/10.3354/cr00936

10.3354/cr00936
109

Poczai P, Varga I, Laos M, Cseh A, Bell N, Valkonen JP, Hyvönen J (2013) Advances in plant gene-targeted and functional markers: a review. Plant Methods 9:6. https://doi.org/10.1186/1746-4811-9-6

10.1186/1746-4811-9-623406322PMC3583794
110

Poorter H, Garnier E (1996) Plant growth analysis: an evaluation of experimental design and computational methods. J Exp Bot 47:1343-1351. https://doi.org/10.1093/jxb/47.9.1343

10.1093/jxb/47.9.1343
111

Poorter H, Sack L (2012) Pitfalls and possibilities in the analysis of biomass allocation patterns in plants. Front Plant Sci 3:259. https://doi.org/10.3389/fpls.2012.00259

10.3389/fpls.2012.0025923227027PMC3514511
112

Prusinkiewicz P (2004) Modeling plant growth and development. Curr Opin Plant Biol 7:79-83. https://doi.org/10.1016/j.pbi.2003.11.007

10.1016/j.pbi.2003.11.007
113

Raikhel NV (2001) Plant physiology: past, present, and future. Plant Physiol 125:1-3. https://doi.org/10.1104/pp.125.1.1

10.1104/pp.125.1.1
114

Ramadan MES (2024) Alleviation of irrigation water salinity impact on the growth and yield of sweet potato by plant growth-promoting rhizobacteria inoculation. Hortic Environ Biotechnol 65:669-677. https://doi.org/10.1007/s13580-024-00598-3

10.1007/s13580-024-00598-3
115

Sansavini S (2008) Horticulture in Europe: from history to innovation. Acta Hortic 817:43-58. https://doi.org/10.17660/ActaHortic.2009.817.2

10.17660/ActaHortic.2009.817.2
116

Schmid B, Harper JL (1985) Clonal growth in grassland perennials: I. density and pattern-dependent competition between plants with different growth forms. J Ecol 73:793-808. https://doi.org/10.2307/2260147

10.2307/2260147
117

Seo TC, An SW, Jang HW, Nam CW, Chun H, Kang TK, Lee SH (2018) An approach to determine the good seedling quality of grafted tomatoes (Solanum lycopersicum) grown in cylindrical paper pot through the relation analysis between DQI and short-term relative growth rate. J Bio Environ Control 27:302-311. https://doi.org/10.12791/KSBEC.2018.27.4.302

10.12791/KSBEC.2018.27.4.302
118

Shah L, Yahya M, Shah SMA, Nadeem M, Ali A, Ali A, Wang J, Riaz MW, Rehman S, et al. (2019) Improving lodging resistance: using wheat and rice as classical examples. Int J Mol Sci 20:4211. https://doi.org/10.3390/ijms20174211

10.3390/ijms2017421131466256PMC6747267
119

Shahid M, Pinelli E, Dumat C (2018) Tracing trends in plant physiology and biochemistry: need of databases from genetic to kingdom level. Plant Physiol Biochem 127:630-635. https://doi.org/10.1016/j.plaphy.2018.04.030

10.1016/j.plaphy.2018.04.030
120

Shin EJ, Lee JH, Nam SY (2023) Changes in growth, visual qualities, and photosynthetic parameters in Peperomia species and cultivars under different color temperatures of white lighting conditions. J Agric Life Environ Sci 35:307-321. https://doi.org/10.22698/jales.20230025

10.22698/jales.20230025
121

Shin EJ, Lee JH, Nam SY (2024) Evaluation of growth, vegetation indices, and photosynthesis of Cichorium intybus L. seedlings as affected by LED light qualities in a closed nursery facility. Hortic Sci Technol 42:350-364. https://doi.org/10.7235/HORT.20240029

10.7235/HORT.20240029
122

Smirnakou S, Ouzounis T, Radoglou KM (2017) Continuous spectrum LEDs promote seedling quality traits and performance of Quercus ithaburensis var. macrolepis. Front Plant Sci 8:188. https://doi.org/10.3389/fpls.2017.00188

10.3389/fpls.2017.0018828261244PMC5306215
123

Song KS, Jeon KS, Kim CH, Yoon JH, Park YB, Kim JJ (2014) Effect of shading level on growth and morphological characteristics of Ligularia fischeri seedling. J Bio Environ Control 23:88-94. https://doi.org/10.12791/KSBEC.2014.23.2.88

10.12791/KSBEC.2014.23.2.88
124

Sour V, Dongsansuk A, Ayutthaya SIN, Ruamrungsri S, Hongpakdee P (2025) Mitigating water stress and enhancing aesthetic quality in off-season potted Curcuma cv. ‘Jasmine Pink’ via potassium silicate under deficit irrigation. Horticulturae 11:856. https://doi.org/10.3390/horticulturae11070856

10.3390/horticulturae11070856
125

Spani F, Locato V, De Gara L (2025) Unveiling nature’s architecture: geometric morphometrics as an analytical tool in plant biology. Plants 14:808. https://doi.org/10.3390/plants14050808

10.3390/plants1405080840094810PMC11901851
126

Srivastava A, Strasser RJ, Govindjee G (1999) Greening of peas: parallel measurements of 77 K emission spectra, OJIP chlorophyll a fluorescence transient, period four oscillation of the initial fluorescence level, delayed light emission, and P700. Photosynthetica 37:365-392. https://doi.org/10.1023/A:1007199408689

10.1023/A:1007199408689
127

Stamford JD, Vialet-Chabrand S, Cameron I, Lawson T (2023) Development of an accurate low cost NDVI imaging system for assessing plant health. Plant Methods 19:9. https://doi.org/10.1186/s13007-023-00981-8

10.1186/s13007-023-00981-836717879PMC9887843
128

Stirbet A, Govindjee G (2011) On the relation between the Kautsky effect (chlorophyll a fluorescence induction) and photosystem II: basics and applications of the OJIP fluorescence transient. J Photochem Photobiol B Biol 104:236-257. https://doi.org/10.1016/j.jphotobiol.2010.12.010

10.1016/j.jphotobiol.2010.12.010
129

Struve DK (2009) Tree establishment: a review of some of the factors affecting transplant survival and establishment. Arboric Urban For 35:10-13. https://doi.org/10.48044/jauf.2009.003

10.48044/jauf.2009.003
130

Suetsugu K, Sugita R, Yoshihara A, Okada H, Akita K, Nagata N, Tanoi K, Kobayashi K (2023) Aerial roots of the leafless epiphytic orchid Taeniophyllum are specialized for performing crassulacean acid metabolism photosynthesis. New Phytol 238:932. https://doi.org/10.1111/nph.18812

10.1111/nph.18812
131

Sun X, Luo S, Luo L, Wang X, Chen X, Lu Y, Shen S, Zhao J, Bonnema G (2018) Genetic analysis of Chinese cabbage reveals correlation between rosette leaf and leafy head variation. Front Plant Sci 9:1455. https://doi.org/10.3389/fpls.2018.01455

10.3389/fpls.2018.0145530337935PMC6180146
132

Tan CW, Huang WJ, Jin XL, Wang JC, Tong L, Wang JH, Guo WS (2012) Monitoring the chlorophyll fluorescence parameter Fv/Fm in compact corn based on different hyperspectral vegetation indices. Spectrosc Spec Anal 32:1287-1291. https://doi.org/10.3964/j.issn.1000-0593(2012)05-1287-05

10.3964/j.issn.1000-0593(2012)05-1287-05
133

Tan CW, Wang DL, Zhou J, Du Y, Luo M, Zhang YJ, Guo WS (2018) Assessment of Fv/Fm absorbed by wheat canopies employing in-situ hyperspectral vegetation indexes. Sci Rep 8:9525. https://doi.org/10.1038/s41598-018-27902-3

10.1038/s41598-018-27902-329934625PMC6015031
134

Tumber-Dávila SJ, Schenk HJ, Du E, Jackson RB (2022) Plant sizes and shapes above and belowground and their interactions with climate. New Phytol 235:1032-1056. https://doi.org/10.1111/nph.18031

10.1111/nph.1803135150454PMC9311740
135

Van Kleunen M, Weber E, Fischer M (2010) A meta-analysis of trait differences between invasive and non-invasive plant species. Ecol Lett 13:235-245. https://doi.org/10.1111/j.1461-0248.2009.01418.x

10.1111/j.1461-0248.2009.01418.x
136

Vonlanthen T, Fuchs Z, Cronje C, Katsir L, Vogt M, George G, Ruckle ME, Hiltbrunner J (2025) Study to develop a value for cultivation and use (VCU) field trial protocol for Cannabis sativa L. flower varieties. Agronomy 15:1338. https://doi.org/10.3390/agronomy15061338

10.3390/agronomy15061338
137

Weiner J (2004) Allocation, plasticity and allometry in plants. Perspect Plant Ecol Evol Syst 6:207-215. https://doi.org/10.1078/1433-8319-00083

10.1078/1433-8319-00083
138

Wells R (1991) Soybean growth response to plant density: relationships among canopy photosynthesis, leaf area, and light interception. Crop Sci 31:755-761. https://doi.org/10.2135/cropsci1991.0011183X003100030044x

10.2135/cropsci1991.0011183X003100030044x
139

Wen S, Shi N, Lu J, Gao Q, Hu W, Cao Z, Lu J, Yang H, Gao Z (2022) Continuous wavelet transform and back propagation neural network for condition monitoring chlorophyll fluorescence parameters Fv/Fm of rice leaves. Agriculture 12:1197. https://doi.org/10.3390/agriculture12081197

10.3390/agriculture12081197
140

Weraduwage SM, Chen J, Anozie FC, Morales A, Weise SE, Sharkey TD (2015) The relationship between leaf area growth and biomass accumulation in Arabidopsis thaliana. Front Plant Sci 6:167. https://doi.org/10.3389/fpls.2015.00167

10.3389/fpls.2015.0016725914696PMC4391269
141

Wi SH, Yeo KH, Seo TC, Lee HJ, Lee H, An S, Jang Y (2024) Influence of light, temperature, and irrigation on grafted cucumber seedlings in biodegradable paper pots. J Bio Environ Control 33:280-293. https://doi.org/10.12791/KSBEC.2024.33.4.280

10.12791/KSBEC.2024.33.4.280
142

Xiao Y, Liu J, Li H, Cao X, Xia X, He Z (2015) Lodging resistance and yield potential of winter wheat: effect of planting density and genotype. Front Agric Sci Eng 2:168-178. https://doi.org/10.15302/J-FASE-2015061

10.15302/J-FASE-2015061
143

Xue J, Gou L, Zhao Y, Yao M, Yao H, Tian J, Zhang W (2016) Effects of light intensity within the canopy on maize lodging. Field Crops Res 188:133-141. https://doi.org/10.1016/j.fcr.2016.01.003

10.1016/j.fcr.2016.01.003
144

Xue J, Su B (2017) Significant remote sensing vegetation indices: a review of developments and applications. J Sen 2017:1353691. https://doi.org/10.1155/2017/1353691

10.1155/2017/1353691
145

Yan Z, Cao X, Bing L, Lin D, Cheng F, Wang K, Qi Y, Yang Y (2025) Assessment of the growth and quality of pepper seedlings under the combinations of daily light integral and nitrogen concentration. Hortic Environ Biotechnol 66:331-346. https://doi.org/10.1007/s13580-024-00655-x

10.1007/s13580-024-00655-x
146

Yang H, Kim Y, Bae Y, Hyeon S, Jang D (2024) Acclimation and hardness enhancement through ultraviolet-B irradiation of fruit vegetable seedlings grown in a plant factory with artificial light. Hortic Environ Biotechnol 65:771-783. https://doi.org/10.1007/s13580-024-00605-7

10.1007/s13580-024-00605-7
147

Yun JH, Jeong HW, Hwang SY, Yu J, Hwang HS, Hwang SJ (2023) Growth of cucumber and tomato seedlings by different light intensities and CO2 concentrations in closed-type plant production system. J Bio Environ Control 32:257-266. https://doi.org/10.12791/KSBEC.2023.32.4.257

10.12791/KSBEC.2023.32.4.257
148

Zhang H, Zhao Y, Zhu JK (2020) Thriving under stress: how plants balance growth and the stress response. Dev Cell 55:529-543. https://doi.org/10.1016/j.devcel.2020.10.012

10.1016/j.devcel.2020.10.012
149

Zhang P, Gu S, Wang Y, Yang R, Yan Y, Zhang S, Sheng D, Cui T, Huang S, et al. (2021) Morphological and mechanical variables associated with lodging in maize (Zea mays L.). Field Crops Res 269:108178. https://doi.org/10.1016/j.fcr.2021.108178

10.1016/j.fcr.2021.108178
150

Zhang X, Zhou Y, Ji Y, Yu M, Li X, Duan J, Wang Y, Gao J, Guo X (2023) Climate factors affect above-belowground biomass allocation in broad-leaved and coniferous forests by regulating soil nutrients. Plants 12:3926. https://doi.org/10.3390/plants12233926

10.3390/plants1223392638068563PMC10708195
151

Zhu M, Lin C, Jiang Z, Yan F, Li Z, Tang X, Yang F, Ding Y, Li W, et al. (2023) Uniconazole enhances lodging resistance by increasing structural carbohydrate and sclerenchyma cell wall thickness of japonica rice (Oryza sativa L.) under shading stress. Environ Exp Bot 206:105145. https://doi.org/10.1016/j.envexpbot.2022.105145

10.1016/j.envexpbot.2022.105145
152

Zolin Y, Popova A, Yudina L, Grebneva K, Abasheva K, Sukhov V, Sukhova E (2025) RGB indices can be used to estimate NDVI, PRI, and Fv/Fm in wheat and pea plants under soil drought and salinization. Plants 14:1284. https://doi.org/10.3390/plants14091284

10.3390/plants1409128440364313PMC12073610
Information
  • Publisher :KOREAN SOCIETY FOR HORTICULTURAL SCIENCE
  • Publisher(Ko) :한국원예학회
  • Journal Title :Horticultural Science and Technology
  • Journal Title(Ko) :원예과학기술지
  • Received Date : 2025-08-29
  • Revised Date : 2026-01-12
  • Accepted Date : 2026-01-14